. "Chordata" . "Grant, Humphrey, and Myers 1997" . "Colostethus atopoglossus" . "Dendrobatidae" . . "Anura" . . "Amphibia" . "Colostethus" . . . "Animalia" . . . "Colostethus atopoglossus Grant, Humphrey, and Myers 1997" . . . . . . .