"Marchantiophyta" . . "Plantae" . . . . "Jungermanniopsida" . . . . "Marsupella" . "R.M. Schust. and Damsh." . . . "Gymnomitriaceae" . . . . "Marsupella spiniloba R.M. Schust. and Damsh." . "Marsupella spiniloba" . "Jungermanniales" .