"Chaparana quadrana"^^ . "Gynandropaa quadranus"^^ . "Feirana quadranus"^^ . . "vto-namespace"^^ . "ATO:0004250"^^ . . . "Paa quadrana"^^ . "Paa quadranus"^^ . "Nanorana quadranus"^^ . "http://amphibiaweb.org/species/7443"^^ . "Nanorana quadrana"^^ . "Nanorana quadranus"^^ . "Rana quadrana [unjustified emendation]"^^ . "Rana quadrana"^^ . "Chaparana quadranus"^^ . "VTO:0003200"^^ . "Rana quadranus"^^ .