. "Heyer & Heyer (2012)" . . "Heyer & Heyer (2012)" . "2012"^^ . . "Heyer, W. R. & Heyer, M. M. 2012. Systematics, distribution, and bibliography of the frog Leptodactylus validus Garman, 1888 (Amphibia: Leptodactylidae). Proceedings of the Biological Society of Washington, 125(3): 276-294." . "2012"^^ .