"Copelatus luctuosus Guignot, 1939" . . . . _:b75965762 . _:b75965762 . "Copelatus luctuosus" . . "Copelatus luctuosus" . . "755319" . "Guignot, 1939" . "Copelatus luctuosus" . . . _:b75965762 . "6001714" . _:b75965762 "GBIF id" . . _:b75965762 "6001714" .